KINGDOM: Protista

KINGDOM: Protista

This kingdom formed a link between kingdom Monera on one hand and other three kingdoms i.e. Plantae, Fungi and Animalia on the other hand. Protistians are ancestors of all multicellular eukaryotes (plants, fungi, animals).

Kingdom: Protista includes-

                                                Protista
                                                    

         (1) Autotrophic     (2) Consumer                (3) Protozoans
                     or
      e.g Dinofagellates    e.g. Cellular slime      e.g.Zooflagellates
           Diatoms                     moulds                      Sarcodines
           Euglenoids                Acellular slime          Sporozoans
                                               moulds                       Ciliates

                                                    

General Characteristics of Protista:-

(1)  Unicellular, some are colonial without much cellular differentiation.

(2) Mostly aquatic organisms.

(3) Cells may possesses cellulosic cell wall.

(4) Flagella and cilia have (9+2) pattern of microtubule organization       consisting of tubulin protein.

(5) Movement of pseudopodia, flagella or cilia where ciliary mode is fastest.

(6) Mode of nutrition may be photosynthetic (holophytic). Holozoic (ingestion), saprobic or parasitic (absorptive). Some heve mixotrophic nutrition (photosynthetic and saprobic) as in Euglena.

(7) Reproduction occurs by asexual and sexual means.

(8) Life cycle is of two types:-

(i) Showing zygotic meiosis
(ii) Showing gametic meiosis    

  1. Photosynthetic in Protists:-

These are populary called protistan algae. Protistan algae constitute the major portion of the phytoplanktons.

  1. Diatoms:-

    Diatoms are golden brown photosynthetic protistis and are called Chrysophytes (including both diatoms and desmids). They are both aquatic and terrestrial. Some are marine. Their important characters are:-

         (i) These are microscopic organisms possessing varying colours.

         (ii) They are basically unicellular, but may form pseudofilament and                       colonies, lacking flagella except in the reproductive stage.  They may               be free floating (phytoplanktonic) , remaining afloat on surface of                       water due to presence of light weight lipids.

         (iii) The cell wall is impregnated with silica to form transparent siliceous                 shell, known as frustules.

         (iv) They are responsible for almost 50% of the total organic matter                       synthesized in the biosphere.

        (v) They are very good indicators of water pollution.

        (vi) Diatoms are predominantly aquatic, inhabiting both marine and                          freshwater environments.

       (vii) They play a critical role in the carbon cycle through photosynthesis,                  producing a significant portion of the world’s oxygen. Diatoms have a                high diversity, with over 100,000 species documented.

       (viii) They reproduce primarily asexually through binary fission, though                        sexual reproduction also occurs under certain conditions.

       (ix) Their fossilized remains form diatomaceous earth, utilized in various                  industrial applications.

       (x) Diatoms are also valuable in paleoclimatology, helping reconstruct                     past climatic conditions based on their preserved silica shells in                         sediment records.

        (xi) Auxospores:- Zygotes formed during sexual reproduction in in                             Diatoms is called Auxospore, common examples of diatoms are                         Triceratium, Melosira, Navicula and Cymbella.

  1. Dinoflagellates:-

    Dinoflagellates are golden brown photosynthetic protists, belonging to class Dinophyceae (Pyrrophyta). They are mainly marine, though few are fresh water forms. Their important characters are:-

        (i) They are unicellular, motile, biflagellated.

       (ii) The is enclosed by a rigid coat called theca or lorica consisting of 2
            to many articulated or sculptured plates of cellulose and pectin,                          hence are called armoured Dinoflagellates.

       (iii) Reserved food is carbohydrate and oils.

      (iv) Reproduction is commonly asexual and occurs through cell division.

      (v) Isogamous and anisogamous sexual reproduction is reported  from                 dinoflagellates e.g Ceratium.

      (vi) Life cycle involves zygotic meiosis (Ceratium, Gymnodinium).
            Gametic meiosis occurs in Noctiluca.

       (vii) They possess two dissimilar flagella, one wrapped around a                                 transverse groove (cingulum) and the other trailing, enabling                               unique motility patterns.

        (viii) Many species exhibit bioluminescence, emitting light when                                  disturbed, contributing to marine phenomena like                                                   “phosphorescent seas.”

         (ix) Some species are known for harmful algal blooms (HABs),
               producing toxins that can impact marine life and human health.

         (x) They are significant in marine ecosystems, contributing to primary                     production and serving as key components in the food web.

protista- dinoflagella

       (iv) The cell bears a single long type flagellum (stichonematic) arising at                 the anterior end. Actually there are two flagella but one of these is                     reduced. The longer flagellum has two branches at the base each                       having its own basal granule. In the area of union of two flagella is                     present a photosensitive paraflagellar body

        (v) Myonemes are oblique but parallel arranged strips in pellicle.                              Euglenoids perform creeping movement of contraction and expan                      with the help of myonemes which is called metaboly or euglenoids                    movement.

      (vi) Stigma or an eye spot is attached to the membrane of the reservoir at              the level of paraflagella body and along with it seems to be involved in              perception of light stimulus. It contains photosensitive red-orange                   pigment called astaxanthin.

      (vii) Single large nucleus lies near the centre of the protoplast.

      (viii) Reserve food material is paramylon.

      (ix) Flagella: They possess one or two flagella emerging from an anterior                 pocket, aiding in motility.

       (x) Pellicle: Instead of a rigid cell wall, euglenoids have a flexible pellicle                composed of protein strips beneath the plasma membrane, allowing                  shape changes.

      (xi) Photosynthesis: Many euglenoids contain chloroplasts with                                 chlorophylls a and b, enabling photosynthesis, though some species                 are heterotrophic

       (xii) Mixotrophy: Euglenoids can switch between autotrophy                                       (photosynthesis) and heterotrophy (ingesting organic matter) based                on environmental conditions

       (xiii) Reproduction: Euglenoids reproduce asexually through longitudinal                 binary fission; sexual reproduction has not been observed.

      (xiv) Habitat: They are predominantly found in freshwater environments,                    though some species inhabit marine and brackish waters.

  1. Euglenoid (Euglena-like):-

    It is a group chlorophyllous flagellate protists. Largest genera being Euglena amongst them.

(i) Euglenoids are unicellular, flagellate protists found in water or damp soil. Majority of them are fresh water organisms found in stagnant water.

(ii) Cell wall is absent but a covering periplast or pellicle is present which is proteinaceous  (elastic) in structure.

(iii) Locomotory organs are flagella.

(iv) The cell bears a single long type flagellum (stichonematic) arising at the anterior end. Actually there are two flagella but one of these is reduced. The longer flagellum has two branches at the base each having its own basal granule. In the area of union of two flagella is present a photosensitive paraflagellar body

(v) Myonemes are oblique but parallel arranged strips in pellicle. Euglenoids perform creeping movement of contraction and expansion with the help of myonemes which is called metaboly or euglenoids movement.

(vi) Stigma or an eye spot is attached to the membrane of the reservoir at the level of paraflagella body and along with it seems to be involved in perception of light stimulus. It contains photosensitive red-orange pigment called astaxanthin.

(vii) Single large nucleus lies near the centre of the protoplast.

(viii) Reserve food material is paramylon.

(ix) Flagella: They possess one or two flagella emerging from an anterior pocket, aiding in motility.

(x) Pellicle: Instead of a rigid cell wall, euglenoids have a flexible pellicle composed of protein strips beneath the plasma membrane, allowing shape changes.

(xi) Photosynthesis: Many euglenoids contain chloroplasts with chlorophylls a and b, enabling photosynthesis, though some species are heterotrophic

(xii) Mixotrophy: Euglenoids can switch between autotrophy (photosynthesis) and heterotrophy (ingesting organic matter) based on environmental conditions

(xiii) Reproduction: Euglenoids reproduce asexually through longitudinal binary fission; sexual reproduction has not been observed.

(xiv) Habitat: They are predominantly found in freshwater environments, though some species inhabit marine and brackish waters.

(2) Slime moulds or consumer-decomposer protists:-

They were included in class myxomycetes of fungi in two-kingdom classification. They were called mycetozoan by DeBary as they are closely related to animals. Mycologists include them in gymnomycota. Because of their nature they are called protistan fungi.

General characteristic of the slime moulds are:-

  1. They usually free-living, creeping over debris like fallen leaves and rotting logs of wood.

        2.  They have naked protoplast, not covered by any cell wall in vegetative               stage.

        3. They lack chlorophyll and have saprobic or phagotrophic mode of                      nutrition.

     4.Slime moulds are unique protists found in moist, decaying organic                     matter.

     5. They exhibit characteristics of both fungi and protozoa but lack chitin in           their cell walls.

    6.Their life cycle includes a motile, amoeboid stage called a plasmodium.

    7. The plasmodium moves to find food, engulfing bacteria and organic                 material through phagocytosis.

   8. Slime moulds play a crucial role as decomposers, breaking down dead             plant material.

    9.This decomposition recycles nutrients back into the ecosystem,                         supporting soil health.

    10.They contribute to plant growth by enriching the soil with nutrients.

   11. Slime moulds can form fruiting bodies that produce spores for                           reproduction.

   12. These spores allow slime moulds to spread to new environments.

   13.Their ability to switch between unicellular and multicellular forms aids in           studying cellular behavior.

 

(3) Protozoans

These are unicellular organism with heterotrophic nutrition.  They are belived to be primitive relatives of animals. There are four groups of protozoans: 

Amoeboid protozoan

Flagellated protozoans

Cilated protozoans

Sporozoans

1. Habitat and habit

   Fresh water, sea                              water or moist soil mostly free living, few parasites.

 

Free living (aquatic) or parasitic

 

Fresh water or marine, few parasite.

 

All endoparasites

2. Locomotory      structure

 Pseudopodia (false feet)

 

 

Flagella

 

 

Cilia

 

 

Absent

3. Special feature

Silica shells in some forms.

 

Rare sexual reproduction with diverse types of association- commensal,symboint, parasitic.

 

Possess definite region of ingestion (gullet) and (egestion).

Infectious spore- like stage is present in life cycle.

4. Example and diseases

 Amoba, Entamoeba (dysentery)

 

 

Trypanosoma (sleeping sickness)

 

 

 Paramoecium

 

 

Plasmodium (most notorious causing malaria)

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